In an alternative Earth in 1939, a deadly alien force known as Neuroi attack nation after nation, destroying military forces. In desperation, several countries band together to create Strike Witches: field aerial combat units of young girls capable of using magic. They soon become the best defence against the invaders.
In 1944, Yoshika Miyafuji is approached by ace witch Mio Sakamoto to join the 501st Joint Fighter Wing. She then must learn what it takes to be a Strike Witch while she struggles to bond with her fellow comrades and take down an enemy she doesn't understand, and doesn't hate.
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